went back to the transformer wiring, traced my connections and double checked my readings at which point I realized that the 275V out from the PS was in the printed limit. Guess I need to go over my shunt board again.
The +275V of the power supply board will go higher if no current is being drawn from the power supply. The MJE5731A collectors will also be much higher than they are supposed to be if no current is being drawn through the circuit.
Yes, the 12AU7 needs to be installed to test the board that lives above the 12AU7. You can slip your black meter probe into the ground post so that you don't have to hold it. The red probe should measure DC voltage on each Kreg pad.
The tabs measure out fine, I'll chalk this up to an opportunity to double check my wiring, found a couple untrimmed leads and 1 connection that I hadn't soldered. Also for future work read then reread the instructions to make certain that I am clear.