The 1V at Kreg and 300V at Breg means the 12AU7 in the middle is sucking down nearly limitless current. The simplest way for this to happen would be a wiring error on the high current C4S boards.
i am not sure what u mean by the OA & OB on the center pc board. the OA connection of the center pc board is a white wire connected to C1. the OB is connected to A6 if thats what u mean .
Resoldering your connections brought one side into proper operation. I would suggest that you also resolder the center PC board and perhaps that will fix the other side.