At this point you'd want to focus on the small board not working properly. You can even unplug the 6080 for now and just run the amp with the 12AU7.
Can you do a DC resistance check between terminal 1 and ground?
Generally when a C4S board fails to function, we see high voltage on the output and resoldering is required. Seeing 0V on the output is more typical for the output being somehow shorted to ground, which suggests debris or wire ends touching as a cause of the problem.